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Just as Gio makes Betty an incredibly romantic offer, Henry pops back into her life with a proposal of his own. Meanwhile, Betty finds herself centerfield at the charity softball game of the year: Mode vs. ELLE Magazine. Can Wilhelmina strike out ELLE’s star player, supermodel Naomi Campbell? In other developments, an orphaned boy from France turns up at Mode with an amazing claim, and Hilda learns some shocking news about Coach Diaz.
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Betty, still missing Henry, fights her attraction to Gio, but later finds herself stuck with him when both volunteer to chaperone Justin’s middle school dance. Meanwhile, Hilda tries to get Coach Diaz (guest star Eddie Cibrian) to notice her; Amanda agrees to do a reality show with her dad, KISS rock legend Gene Simmons; and Daniel feels the heat when Wilhelmina makes her triumphant return to Mode amid a media frenzy.
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Betty switches gears with rival Charlie (guest star Jayma Mays) and tries to befriend her by throwing a baby shower, a party that no one will ever forget. Meanwhile, Hilda confronts Justin’s PE teacher (guest star Eddie Cibrian as Coach Diaz) when she learns Justin is flunking gym; Daniel unravels after his break-up with Renee, and Christina finally learns who fathered the baby she’s carrying for Wilhelmina.
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An increasingly delusional Renee (guest star Gabrielle Union) becomes convinced that Betty is in love with Daniel and therefore is her rival for his affections. Meanwhile, Hilda’s new client at the beauty shop turns out to be ex-neighbor and lifetime nemesis Gina Gambarro (guest star Ava Gaudet), who returns to Queens to rub her fabulous life and wealthy doctor husband in Hilda’s face.
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It is Betty’s 24th birthday and she wants it to be perfect. Unfortunately, her plan of going out with Henry is disrupted when Charlie returns to New York to be with him. Gio kindly gives Betty a dream birthday. Meanwhile, Daniel is still dating Renee, whom Wilhelmina still unsuccessfully tries to keep Daniel away from.
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Encouraged by both Gio and Henry, Betty accepts her first writing assignment and ends up researching picking up artist techniques. Daniel is fascinated with a new woman, Renee, who is more related to him than he is aware. Amanda continues her search for her father. Claire is determined to start a new magazine against the will of her children. Henry tells Betty how he feels about Gio and Betty has to make a decision about it.
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While the whole Meade family is involved in Claire’s trial on Fey Sommer’s murder, Betty comes upon a perfume that makes her go hyper-mode and drives both Henry and Gio crazy about her. After violently breaking into Gio’s deli and ending in a hospital, Betty and her family discover the deadly truth about the mysterious perfume. Amanda has a moral dilemma regarding the past of her late mother and Wilhelmina, with the help of Marc, will try the impossible to make Christina agree to join her in her mischievous plans.
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Amanda visits a psychic (guest star Annie Potts) for insight into her dad’s identity. Meanwhile, disturbed about the message Mode is sending to Justin and his classmates after they visit the magazine, Betty pushes Daniel to use “real” models during fashion week; and Wilhelmina gets some unexpected news about her latest plans for the Meade dynasty.
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Unable to decide who their father left in charge of the Meade Empire, Daniel and Alexis settle the matter with a paint ball fight inside MODE offices. Trying to spice her love life, Betty drags Henry into a double date with romantic assertive Gio and her very outgoing sister, Hilda, at a lively reggaeton club. Wilhelmina faces financial problems in new launched Slater Magazine and tries to improve her public image ending in a total media disaster as she ran over Betty White by mistake. Amanda and Nick finally settle their differences.
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A Wilhelmina-induced crisis at Mode finds everyone scrambling to recreate an issue on deadline; Daniel and Betty try to break the wild-child cover girl out of rehab for a reshoot; Alexis tries to charm the printer into extending their time.