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In the wake of the disastrous Meade/Slater nuptials, Wilhelmina plots her next moves and Betty tries to make things right with Daniel. Meanwhile, Henry uses his charms to get the Mode weekend receptionist to let him in after hours so he can help Betty get something of vital importance.
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As Wilhelmina prepares for her big day, all is not quiet on the Mode front when she realizes her fabulous maid of honor, Victoria Beckham (guest starring as herself), is stealing her thunder. Meanwhile, Betty is wracked with guilt about her secret pact with Wilhelmina and about not telling Daniel that his future step-mom is dallying with her bodyguard (guest star Rick Fox). At the Suarez house, Ignacio prepares for his big day – becoming a U.S. citizen – but when Betty moves in with new love Henry, it puts a major strain on father/daughter relations.
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Betty and Henry continue their illicit romance and plan a secret date to “Wicked,” but things don’t go as planned when Daniel, thinking she’s dating Gio (Freddy Rodriguez), gifts her with tickets for the two of them instead – tickets which Gio refuses to give up. Meanwhile, Daniel woos an important advertiser (guest star Marlo Thomas), a lady with a penchant for younger men.
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Much to Henry’s distress, Betty tries internet dating and gets asked out on a bowling date. Meanwhile Marc finds himself attracted to someone who isn’t his type, chubby but charming photographer Cliff. On the home front, Daniel and Alexis steel themselves for a “family” dinner with step-mom-to-be Wilhelmina, and Hilda, lost in her own world, doesn’t notice that Justin is ditching school and running with the wrong crowd.
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When Mode is forced to abandon its big Christmas photo shoot because another magazine has copied the idea, Betty forms an uneasy alliance with Marc and Amanda. The three fear they may have leaked the idea to someone at the rival mag (guest star Nicholas Gonzalez, “The OC”) during happy hour… and that their jobs will be history if anyone finds out. Meanwhile, Daniel and Wilhelmina reluctantly work together to come up with a new photo spread as Mode’s deadline looms, and Betty’s nephew, Justin, is oblivious to the turmoil as he spends a day at the magazine for a class project.
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Betty tries to avoid thinking about her problems with boyfriend Henry and focuses on helping Daniel in the annual Mode benefit event but suddenly encounters the new sandwich cart employee, Gio, who both fascinates her and challenges her to follow her dreams. Henry has to face ex-girlfriend Charlie after finding shocking news about her pregnancy. Wilhelmina finds new ways to drag Alexis back to her side. Ignacio is in Mexico facing a deadly threat from his past while Marc and Amanda plan new ways to publicly promote Amanda as Fey Sommer’s daughter.
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Bradford tries to change the Meade family history by taking advantage of Alexis’s amnesia; Betty suspects Wilhelmina’s new bodyguard is doing a lot more than protecting her; Ignacio is in for a surprise from his past; Amanda receives a portion of Fey’s inheritance…and the expenses and fees that come with it.
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After so many sad events in Betty’s life, she can’t seem to get her life into perspective. She denies that she is wrecked by the fact that Henry is leaving. Amanda on the other hand is shocked after discovering who her real mother is. Of course, Wilhelmina is right around the corner arranging to take advantage of everything going on around her