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Betty and Henry’s romance heats up. Meanwhile, Wilhelmina makes great strides in her ambitious power play, and Daniel descends further in a downward spiral. The Suarez family trip to Mexico takes a dangerous turn, as old ghosts create new problems. Explosive secrets will be revealed, and everything will be tied together by a beloved Broadway musical.
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Betty and family head to Mexico to get Ignacio his visa, Wilhelmina puts on a show for Amanda’s new boyfriend, Tavares, and Alexis’ ex-girlfriend comes to visit.
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At a medieval-themed restaurant where Mode assistants celebrate Secretaries’ — er, Administrative Assistants’ Day, Henry battles to defend Betty’s honor (which leaves girlfriend Charlie less than thrilled). Betty convinces Daniel that he has a sex addiction and needs help.
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Betty, fed up with Daniel, Christina and Henry and how they’ve all disappointed her, does her best to avoid all three. Meanwhile, love’s all around at Mode, as Alexis gets asked out on her first date as a woman; Amanda begins a flirtation with a designer (guest star Mykel Shannon Jenkins) who she assumes is gay; Wilhelmina struggles to deal with the tireless advances of her latest conquest, and Hilda gets a surprise from Santos (guest star Kevin Alejandro). Meanwhile, Daniel goes it alone in dealing with the aftermath of his one-time hook-up with Petra.
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In Punch Out, Daniel meets a very young model, Betty and Christina go clubbing, Wilhelmina seduces Bradford, Alexis looks for a secret room, and Constance tries to marry Ignacio.
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Marc, promising to help Betty with inside info for Daniel, convinces her to pretend to be his “girlfriend” when his suspicious mom, Mrs. Weiner (Tony winner Patti LuPone), arrives in town. Things go from bad to worse when Mrs. Weiner invites herself to dinner at the Suarez home and Betty’s whole family has to play along with the charade. Meanwhile, Daniel and Alexis struggle to work together after their mom – who actually owns MODE – names them co-editors-in-chief of the magazine.
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Daniel keeps secret from Alexis the fact that he is the one who hired Grace to defend their father, something which Wilhelmina uses to bring the siblings apart. Meanwhile, when invited by Henry to Charlie’s birthday party, a desperate Betty asks Dr. Farkas, her orthodontist, to be her date. Finally, Amanda catches the eye of an unusual designer who claims that she is his muse.
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Family matters force Daniel to reconnect with Grace Chin (guest star Lucy Liu in the first of a two-episode arc), a girl he once stood up for a date in college. She’s now known as “The Chin,” NYC’s most powerful lawyer. Meanwhile, Betty makes a new friend, Charlie (guest star Jayma Mays), a likable out-of-towner she meets in the Mode cafeteria, only to find out Charlie is Henry’s girlfriend. And a blizzard descends on the city and derails plans, as Christina, trapped in the office with Marc, struggles to put together a gown for Sarah Jessica Parker; Wilhelmina and Alexis get stranded in a sports bar; and Hilda, Santos and Justin get stuck on the subway on the way to see Justin’s favorite musical, “Hairspray.”
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In Brothers, there’s a power struggle at Mode when Alexis returns, Henry asks Betty out on a date to see Wicked, and Justin gets into a fight at school.
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It’s Fashion Week in New York and Betty is hustling to ensure “Mode’s” runway show goes off without a hitch. To Betty’s dismay, Daniel offers Hilda a job helping out with the show. Wilhelmina is introduced to a new “cosmetic technique,” as Christina prepares for the debut of her very own designs. Marc and Amanda resume their scheming ways in a mission to steal what will be the season’s hottest fashion item. Meanwhile, Daniel is moved by his father’s decision to hand over the reigns of Meade Publications, when a visit from an old friend coaxes him into his old ways. But nothing can prepare them for the frenzy that ensues when the “Mode” show is abruptly halted by a shocking revelation.